Infiltrating the Predacon base, Dinobot steals the two Golden Disks from Waspinator, but not before the two agree that with only one moon above, there is now proof that this planet is indeed Earth! Megatron travels to a distant, sparse forest where two stasis pods have crashed, and welcomes to his side two new Predacons: Quickstrike and Silverbolt. His army bolstered, Megatron prepares an all-out attack on the undermanned Maximal base.
With Rhinox unconscious and Tigatron and Airrazor off to scout for fallen stasis pods, only three Maximals are left to defend the Axalon. The Predacons attack while Blackarachnia secretly downloads Megatron's back-up files of the two Golden Disks. Just as the battle looks most hopeless, Silverbolt defects to the Maximals, and a surprise from above turns the tide.
Optimus Primal is back and better than ever in his new TransMetal body. Megatron dispatches Blackarachnia, Tarantulas, and Quickstrike to a nearby cave to build a refueling station. Rattrap and Silverbolt track the trio, and when a fight ensues, the cave is destroyed and the Predacons retreat. Megatron is furious at their failure, but little does he realize Tarantulas' plans for the now-forgotten cave.
What does honor mean to a noble and complicated warrior such as Dinobot? And is he willing to betray that honor in order to tip the scales of the Beast Wars? Dinobot hands the first Golden Disk over to Megatron, betraying the Maximals, and joining the ranks of the Predacons.
The Maximals detect another alien device far away from the Axalon and investigate. Spying a mammoth plant/structure, the Maximal duo are attacked by strange creeping vines, converted into energy and beamed into space. The Predacons arrive to seize the device for themselves, but it may prove too powerful for them.
Megatron boards the Metalhunter, the activated plant/structure at the center of the alien site. While Tarantulas theorizes about its connection to the aliens, TransMetal technology, and the Beast Wars at large, Megatron sets his sights on the Maximals with this floating fortress at his disposal.
What is the mystery of Protoform X? In the jungle there lies a single, oversized stasis pod with a seemingly dead TransMetal inside. Optimus Primal and Megatron both know that this prize must be theirs, but can any Maximal or Predacon stand against the might of the deranged Transformer contained within that pod?
Using his presence on Prehistoric Earth strategically, Megatron employs the Golden Disk to pinpoint how he will change the future to rewrite the timeline and ensure Predacon rule. Dinobot learns of this, and gives his all to stop the Predacons from incinerating the valley from which humans are destined to evolve.
Silverbolt and Rattrap discover another stasis pod, but this one contains a strange, malformed Transformer. Unable to the speak clearly and lacking the specs of a scientist or warrior, the "Transmutate" is both puzzled and puzzling. Silverbolt sees in this simple machine a kindred spirit, an innocent. Predacon Rampage sees something as well: Another one-of-a-kind freak, as alone as he is.
Rhinox tracks the Transwarp Wave from "Aftermath," announcing that Cybertron will soon know of the Axalon's location in space and time. But on a far away Cybertronian space station, the Tripredacus Council deflects the wave and dispatches an agent to Prehistoric Earth. The agent arrives and offers to help, presenting himself to the Maximals as Ravage, formerly of the Decepticon Army! But first, Megatron sends his own troops on a secret mission.
Optimus Primal is concerned that Silverbolt's feelings for the Predacon Blackarachnia are interfering with his duty as a Maximal. Aboard his ship, Ravage taunts the captive Megatron, but Megatron reveals his true intentions for stealing the Golden Disk and its connection with the original Generation One Megatron! Meanwhile, Silverbolt and Blackarachnia arrive at a familiar volcano.
Megatron's grand scheme comes to fruition as the Maximals arrive too late at the site of the crash-landed Autobot Ark. Can Optimus Primal, Rhinox, Cheetor, Silverbolt, and Rattrap stop this plan from undoing the very future of the Transformers?